Inocybe with knobbly spores for ID assistance

[RobSutton]

Found this rather lovely (to my mind anyway) specimen growing in Polytrichum underneath birch at Malham Tarn.



Did as much microscopy as I could, spores:



Cheilocystidia - with crystal like encrustation:



and even what I think are thick walled Pleurocystidia, this one with the same crystal like encrustations:



... and this one without as far as I could see:



I got it down to Inocybe lanuginosa var. ovatocystis using FoS but wasn't convinced by the shape of the cheilocystidia which seemed to match Inocybe leptophylla better but according to FoS in that species they should have thin walled cheilocystidia, no crystals and no pleurocystidia.

I suspect I'm pretty close to getting this one but could do with a nudge in the right direction, or maybe I need a big shove towards something else completely different.

Thanks in advance for any suggestions.

PS I wonder if they've got the photos in FoS for these two the wrong way round, the "Description of the collection examined and illustrated" seems to match the wrong species if you see what I mean.

[mollisia]

Hello,

very close, but it is Inocybe lanuginosa var. lanuginosa.
The var. ovatocystis has only ballon- to pear-shaped Cheilocystidia. But the Cheilos yiou show are much to slender, at least some of them.
I. leptophylla has no pleurocystidia at all, and the Cheilos are thinwalled and without cristals. It is the only knob-spored Inocybe which has non-metuloid cheilos and missing pleuros. If it where a smooth spored species it would key out in the rimosa-group ....

Those three species or varieties are inseperable macroscopically, so the pictures in FoS suite all three for all three taxa ...

best regards,
Andreas

[RobSutton]

Thanks once again Andreas - I can't think why I discounted this species, especially as it's so close to the other two.